CoptoclavidaeDavid R. Maddison
This Mesozoic family consists of medium-sized to large aquatic beetles with divided compound eyes. Larvae have raptorial front legs. The subfamily Necronectinae is found in the Middle to Late Jurassic of Asia and Europe, and Early Cretaceous of northern Africa, Coptoclavinae in the Early Cretaceous of Asia, and Charonoscaphinae in the Late Jurassic of Asia. See Ponomarenko (1977, 1992) for further details.
The following pictures are of larvae of the genus Coptoclava, from the Early Cretaceous of the Shandong Province of China.
Ponomarenko, A.G. 1977. Suborder Adephaga. Pp. 17-104 in Arnold'di, L.V., V.V. Zherikhin, L.M. Nikritin, and A.G. Ponomarenko (eds.), Mezozoiskie Zhestkokrylye. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta, Vol. 161. Nauka Publishers, Moscow.
Ponomarenko, A.G. 1992. Suborder Adephaga. Pp. 19-142 in Arnold'di, L.V., V.V. Zherikhin, L.M. Nikritin, and A.G. Ponomarenko (eds.), Mesozoic Coleoptera. Smithsonian Institution Libraries, Washington, D.C. (English translation of Ponomarenko, 1977.)
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David R. Maddison
Oregon State University
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Maddison, David R. 1995. Coptoclavidae. Version 01 January 1995 (under construction). http://tolweb.org/Coptoclavidae/8896/1995.01.01 in The Tree of Life Web Project, http://tolweb.org/