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Containing group: Athenaria


Carlgren (1949) referred Halcampogeton and Synhalcampella, which previously had been placed in the Halcampoididae to the Edwardsiidae because they have only 8 perfect mesenteries.


Athenaria with elongate, vermiform body usually divisible at least into two regions, a long scapus provided with a cuticle and a short upper scapulus. Often there is also a rounded, naked physa at the aboral end and a very short, thin capitulum immediately below the tentacles. No sphincter or acontia. Mesenteries divisible into 8 macro- and at least 4 microcnemes. Of the macrocnemes there are two pairs of directives and four lateral mesenteries, two on each side, whose retractors face the ventral directives. Retractors diffuse to strongly restricted. Parietal muscles always distinct.


Carlgren, O. 1949. A Survey of the Ptychodactiaria, Corallimorpharia and Actiniaria. Kungl. Svenska Vetenskapsakadamiens Handlingar, series 4, volume 1, number 1.

Title Illustrations
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Nematostella vectensis with egg masses.
Photo copyright © 2000, Ron Ates.
Permission by Prof. C. Hand to take this picture in the Bodega Marine Lab, University of California is gratefully acknowledged.

Copyright © 2000 Ron Ates
About This Page
The information provided on this page is based on Oscar Carlgren's 1949 catalog.
Copyright © 1949 Swedish Academy of Sciences.

Please note that Carlgren's text contains a number of errors, and much of the information is now out of date. An update of the catalog is currently under preparation in Daphne Fautin's laboratory, and the results of this work will be incorporated in future versions of this page.

Keyboarding of Carlgren's catalog was done as part of a project to create an electronic database of the sea anemones of the world, funded by NSF Grant DEB9521819, awarded to Daphne G. Fautin. This grant is in the program Partnerships to Enhance Expertise in Taxonomy (PEET). Susanne Hauswaldt, Katherine Pearson, and April Wakefield-Pagels contributed to the keyboarding effort.

Correspondence regarding this page should be directed to Daphne G. Fautin at

All Rights Reserved.

Citing this page:

Tree of Life Web Project. 2000. Edwardsiidae. Version 01 January 2000 (temporary). http://tolweb.org/Edwardsiidae/17710/2000.01.01 in The Tree of Life Web Project, http://tolweb.org/

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