Scarabaeinae
- Alloscelus
- Ateuchus
- Canthon
- Copris
- Coptodactylus
- Deltochilum
- Dichotomius
- Eucranium
- Eurysternus
- Gymnopleurus
- Heliocopris
- Melanocanthon
- Oniticellus
- Onitis
- Onthophagus
- Phanaeus
- Pinotus
- Scarabaeus
- Sisyphus
Note: this taxon list is still under construction. It does not yet contain all known Scarabaeinae subgroups.
References
Forgie, S. A., T. K. Philips, and C. H. Scholtz. 2005. Evolution of the Scarabaeini (Scarabaeidae: Scarabaeinae). Systematic Entomology 30(1):60-96.
Gill, B. D. 2002. Scarabaeidae Latreille 1802: Scarabaeinae Latreille 1802. Chapter 34II in American Beetles. Volume 2. R. H. Arnett, M. C. Thomas, P. E. Skelley, and J. H. Frank, eds. CRC Press, Boca Raton, 861 pp.
Masumoto, K., K.-M. Chen, and T. Ochi. 2004. A revisional study of the Taiwanese Scarabaeinae (Coloptera, Scarabaeidae). Elytra 32(1):125-131.
Monaghan, M. T., D. J.G. Inward, T. Hunt and A. P. Vogler. 2007. A molecular phylogenetic analysis of the Scarabaeinae (dung beetles). Molecular Phylogenetics and Evolution 45(2):674-692.
Philips, T. K., W. D. Edmonds, and C. H. Scholtz. 2004. A phylogenetic analysis of the New World tribe Phanaeini (Coleoptera: Scarabaeidae: Scarabaeinae): Hypotheses on relationships and origins. Insect Systematics and Evolution 35(1):43-63.
Philips, T. K., E. Pretorius, and C. H. Scholtz. 2004. A phylogenetic analysis of dung beetles (Scarabaeinae: Scarabaeidae): unrolling an evolutionary history. Invertebrate Systematics 18:53-88.
Philips, T. K., C. H. Scholtz, and F. C. Ocampo. 2002. A phylogenetic analysis of the Eucraniini (Coleoptera: Scarabaeidae: Scarabaeinae). Insect Systematic and Evolution 33(3):241-252.
Wilson, C. J. and R. B. Angus. 2005. A chromosomal analysis of 21 species of Oniticellini and Onthophagini (Coleoptera: Scarabaeidae). Tijdschrift voor Entomologie 148:63-76.
Title Illustrations

| Scientific Name | Melanocanthon nigricornis |
|---|---|
| Location | Deep Fork Wildlife Refuge, Okmulgee County, Oklahoma, USA |
| Specimen Condition | Live Specimen |
| Size | approx. 10mm |
| Source Collection | BugGuide.Net |
| ToL Image Use |
This media file is licensed under the Creative Commons Attribution-NonCommercial-NoDerivs License - Version 2.0.
|
| Copyright | © 2007 Charles Schurch Lewallen |
| Scientific Name | Phanaeus igneus |
|---|---|
| Location | Belleview, Marion County, Florida, USA |
| Sex | Female |
| Size | 20mm |
| Source Collection | BugGuide.Net |
| ToL Image Use |
This media file is licensed under the Creative Commons Attribution-NonCommercial-NoDerivs License - Version 2.0.
|
| Copyright | © 2006 Jeff Hollenbeck |
| Scientific Name | Canthon imitator |
|---|---|
| Location | Rancho La Lomitas, Starr County, Texas, USA |
| Specimen Condition | Live Specimen |
| Behavior | Rolling dung near a stable |
| Size | 14 mm |
| Source Collection | Bugwood Network/Forestry Images |
| ToL Image Use |
This media file is licensed under the Creative Commons Attribution-NonCommercial-NoDerivs License - Version 2.5.
|
| Copyright |
© Patrick Coin
|
About This Page
Page copyright © 2007
All Rights Reserved.
- First online 04 May 2007
- Content changed 04 May 2007
Citing this page:
Tree of Life Web Project. 2007. Scarabaeinae. Version 04 May 2007 (temporary). http://tolweb.org/Scarabaeinae/104980/2007.05.04 in The Tree of Life Web Project, http://tolweb.org/





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