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Bark lice

Emilie Bess and Kevin P. Johnson
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taxon links [up-->]Atropetae [up-->]Prionoglaridetae [up-->]Psyllipsocetae [down<--]Psocodea Interpreting the tree
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The root of the current tree connects the organisms featured in this tree to their containing group and the rest of the Tree of Life. The basal branching point in the tree represents the ancestor of the other groups in the tree. This ancestor diversified over time into several descendent subgroups, which are represented as internal nodes and terminal taxa to the right.

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Containing group: Psocodea


The suborder Trogiomorpha includes the families Lepidopsocidae, Psoquillidae, and Trogiidae, comprising more than 300 species in 37 genera.  Trogiomorphs are distributed worldwide with the greatest diversity in the tropics.



General characters:

Discussion of Phylogenetic Relationships

Monophyly of the suborder Trogiomorpha is strongly supported by the unique characters of the female genitalia (described above) and by molecular analysis including all three families (17 species, 15 genera; genes 18S & Histone3 nDNA, 16S rDNA; Yoshizawa et al. 2006).

Other Names for Trogiomorpha


Lienhard, C. and C.N. Smithers. 2002. Psocoptera (Insecta) World Catalogue and Bibliography. Muséum d'Histoire Naturelle, Geneva, Switzerland.

Mockford, E.L. 1993. North American Psocoptera (Insecta). Gainesville, Florida: Sandhill Crane Press.

Smithers, C.N. 1996. Psocoptera. Pp. 1-80, 363-372 (Index) in Wells A. (ed.) Zoological Catalogue of Australia. Vol. 26. Psocoptera, Phthiraptera, Thysanoptera. Melbourne: CSIRO Publishing, Australia.

Smithers, C.N. 1972. The classification and phylogeny of the Psocoptera. Memoirs of the Australian Museum 14: 1–349.

Yoshizawa, K., C. Lienhard, and K.P. Johnson. 2006. Molecular systematics of the suborder Trogiomorpha (Insecta: Psocodea: ‘Psocoptera’). Zoological Journal of the Linnean Society 146: 287–299.

Title Illustrations
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Scientific Name Echmepteryx hageni
Location Dixville, N.H.
Comments 10/9/2007
Specimen Condition Live Specimen
Life Cycle Stage adult
View dorsal
Copyright © Tom Murray
Scientific Name Cerobasis
Location San Pedro, Los Angeles County, California, USA
Size 1.84 mm
Source Photo#50137
Source Collection BugGuide.Net
Image Use creative commons This media file is licensed under the Creative Commons Attribution-NonCommercial-NoDerivs License - Version 1.0.
Copyright © David Ellsworth
Scientific Name Dorypteryx longipennis
Specimen Condition Live Specimen
Identified By Charles Lienhard
Sex Female
Life Cycle Stage adult
Copyright © 2009
About This Page

Emilie Bess
Illinois Natural History Survey, Champaign, Illinois, USA

Kevin P. Johnson
Illinois Natural History Survey, Champaign, Illinois, USA

Correspondence regarding this page should be directed to Emilie Bess at and Kevin P. Johnson at

All Rights Reserved.

Citing this page:

Bess, Emilie and Kevin P. Johnson. 2009. Trogiomorpha. Bark lice. Version 18 March 2009 (under construction). http://tolweb.org/Trogiomorpha/14450/2009.03.18 in The Tree of Life Web Project, http://tolweb.org/

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