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Emilie Bess and Kevin P. Johnson
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taxon links [up-->]Dasydemellidae [up-->]Amphipsocidae [up-->]Asiopsocoidea [up-->]Caeciliusidae [up-->]Stenopsocidae Not Monophyletic[down<--]Psocomorpha Interpreting the tree
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Containing group: Psocomorpha


Infraorder Caeciliusetae includes families Amphipsocidae, Asiopsocidae, Caeciliusidae,  Dasydemellidae, and Stenopsocidae, comprising more than 60 genera and approximately 900 species.  These bark lice are distributed worldwide; about half of the species are described from Asia.  They are small to large in size, about 2–10 mm in length.  Most live on live foliage, but species of Asiopsocidae live on bark.



General Characters

Discussion of Phylogenetic Relationships

Monophyly of Infraorder Caeciliusetae is strongly supported by morphological and molecular data.  Molecular analysis included one nuclear gene (18S) and three mitochondrial genes (16S, 12S, COI; Johnson et al. 2004, Johnson & Mockford 2003).  Morphological data support monophyly of the infraorder based on the synapomorphic characters listed under Characteristics (Yoshizawa 2002).


Johnson, K. P. & E. L. Mockford. 2003. Molecular systematics of Psocomorpha (Psocoptera). Systematic Entomology 28: 409-40

Johnson, K. P., K. Yoshizawa, and V. S. Smith. 2004. Multiple origins of parasitism in lice. Proceeding of the Royal Society of London B 271: 1771-1776.

Lienhard, C. and C. N Smithers. 2002. Psocoptera (Insecta) World Catalogue and Bibliography. Muséum d'Histoire Naturelle, Geneva, Switzerland.

Mockford, E. L. 1993. North American Psocoptera (Insecta). Gainesville, Florida: Sandhill Crane Press.

Mockford, E. L. 2005. Order Psocoptera: Psocids. Pp. 341-355 in C. A. Triplehorn and n. F. Johnson (eds.) Borror and DeLong's Introduction to the Study of Insects. Belmont, California: Thomson Publishing.

Smithers, C. N. 1996. Psocoptera. Pp. 1-80, 363-372 (Index) in A. Wells (ed.) Zoological Catalogue of Australia. Vol. 26. Psocoptera, Phthiraptera, Thysanoptera. Melbourne: CSIRO Publishing, Australia.

Yoshizawa, K. 2002. Phylogeny and higher classification of suborder Psocomorpha (Insecta: Psocodea:'Psocoptera'). Zoological Journal of the Linnean Society 136: 371-400.

Title Illustrations
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Scientific Name Graphopsocus cruciatus
Location Dunstable, Ma.
Comments 10/14/2007
Specimen Condition Live Specimen
Sex m
Life Cycle Stage adult
View lateral
Copyright © Tom Murray
Scientific Name Valenzuela flavidus
Location Groton, Ma.
Comments 9/22/2007
Specimen Condition Live Specimen
Life Cycle Stage adult
View frontal
Copyright © Tom Murray
Scientific Name Matsumuraiella radiopicta
Location Japan
Specimen Condition Live Specimen
Identified By K. Yoshizawa
Life Cycle Stage adult
View lateral
Size 3mm
Copyright © 2006 Kazunori Yoshizawa
About This Page

Emilie Bess
Illinois Natural History Survey, Champaign, Illinois, USA

Kevin P. Johnson
Illinois Natural History Survey, Champaign, Illinois, USA

Correspondence regarding this page should be directed to Emilie Bess at and Kevin P. Johnson at

All Rights Reserved.

Citing this page:

Bess, Emilie and Kevin P. Johnson. 2009. Caeciliusetae. Version 25 March 2009 (under construction). http://tolweb.org/Caeciliusetae/30254/2009.03.25 in The Tree of Life Web Project, http://tolweb.org/

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