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Notholaena copelandii C.C. Hall 1950

Copeland's Cloakfern

Carl Rothfels
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Containing group: core Notholaena

Introduction

Often treated as N. candida var. copelandii, Notholaena copelandii differs from N. candida in morphology (Hall, 1950), farina chemistry (Wollenweber, 1984; Wollenweber et. al 2001), geographic range, and plastid DNA sequences (Rothfels et al., 2008). I thus follow Windham (1993a) and Mickel and Smith (2004) in treating them as distinct species.

Notholaena copelandii
is a species of southern Texas and northeastern Mexico, where it occurs on limestone cliffs and outcrops.

Characteristics

The morphological characters distinguishing N. copelandii from N. candida include: basal pinnae shape (a single exaggerated basioscopic pinnule on N. copelandii; pinnules gradually tapering in length in N. candida); leaf tip shape (abruptly tapered vs. gradually tapered); and rhizome scales (ciliate vs. slightly toothed). Notholaena copelandii is also similar to N. jacalenis, but the later species has much narrower leaves.

Other Names for Notholaena copelandii C.C. Hall 1950

References

Giauque, M. F. A. 1949. Wax glands and prothallia. American Fern Journal 39:33-35.

Hall, C. C. 1950. Notholaena copelandii, a newly recognized species of the Texano-Mexican region. American Fern Journal 40:178--187.

Mickel, J. T., and A. R. Smith. 2004. The Pteridophytes of Mexico. The New York Botanical Garden Press, New York.

NatureServe. 2008. NatureServe Explorer, Arlington, Virginia. www.natureserve.org/explorer/

Rothfels, C. J., M. D. Windham, A. L. Grusz, G. J. Gastony, and K. M. Pryer. 2008. Toward a monophyletic Notholaena (Pteridaceae): Resolving patterns of evolutionary convergence in xeric-adapted ferns Taxon 57:712-724.

Tryon, R. M. 1956. A revision of the American species of Notholaena. Contributions from the Gray Herbarium 179:1-106.

Windham, M. D. 1993a. Notholaena. Pages 143--149 in Flora of North America (Flora of North American Editorial Committee, ed.) Oxford University Press, New York.

Windham, M. D., and G. Yatskievych. 2003. Chromosome studies of cheilanthoid ferns (Pteridaceae: Cheilanthoideae) from the western United States and Mexico. American Journal of Botany 90:1788-1800.

Wollenweber, E. 1984. Exudate flavonoids of Mexican ferns as chemotaxonomic markers. Rev. Latinoamer. Quim. 15:3-11.

Wollenweber, E., M. Drr, and J. F. Stevens. 2001. A dihydroflavonol with taxonomic significance from the fern Notholaena sulphurea. Zeitschrift fr Naturforschung 56:499-502.

Title Illustrations
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Scientific Name Notholaena copelandii
Location USA, Texas
Specimen Condition Live Specimen
Identified By C.J.Rothfels
Image Use creative commons This media file is licensed under the Creative Commons Attribution-NonCommercial License - Version 3.0.
Copyright © 2008 Carl Rothfels
Scientific Name Notholaena copelandii
Location USA, Texas
Specimen Condition Live Specimen
Identified By C.J.Rothfels
Life Cycle Stage sporophyte
Image Use creative commons This media file is licensed under the Creative Commons Attribution-NonCommercial License - Version 3.0.
Copyright © 2008 Carl Rothfels
About This Page

Carl Rothfels
Duke University, Durham, North Carolina, USA

Correspondence regarding this page should be directed to Carl Rothfels at

Page: Tree of Life Notholaena copelandii C.C. Hall 1950. Copeland's Cloakfern. Authored by Carl Rothfels. The TEXT of this page is licensed under the Creative Commons Attribution-NonCommercial License - Version 3.0. Note that images and other media featured on this page are each governed by their own license, and they may or may not be available for reuse. Click on an image or a media link to access the media data window, which provides the relevant licensing information. For the general terms and conditions of ToL material reuse and redistribution, please see the Tree of Life Copyright Policies.

Citing this page:

Rothfels, Carl. 2008. Notholaena copelandii C.C. Hall 1950. Copeland's Cloakfern. Version 23 December 2008 (under construction). http://tolweb.org/Notholaena_copelandii/133605/2008.12.23 in The Tree of Life Web Project, http://tolweb.org/

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