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Emilie Bess and Kevin P. Johnson
Containing group: Atropetae


The family Psoquillidae includes 5 genera and 27 families distributed worldwide.  Many of the seven North American species were introduced at ports of entry; others have been found in warm climates of Florida, Texas, and California.



General Characters

Discussion of Phylogenetic Relationships

The monophyly of the family Psoquillidae is supported by one character in females: the presence of accessory bodies at the opening of spermatheca (Mockford 1993).  The monophyly of Psoquillidae has not been studied with molecular techniques.


Lienhard, C. and C.N. Smithers. 2002. Psocoptera (Insecta) World Catalogue and Bibliography. Muséum d'Histoire Naturelle, Geneva, Switzerland.

Mockford, E.L. 1993. North American Psocoptera (Insecta). Gainesville, Florida: Sandhill Crane Press.

Smithers, C.N. 1996. Psocoptera. Pp. 1-80, 363-372 (Index) in Wells A. (ed.) Zoological Catalogue of Australia. Vol. 26. Psocoptera, Phthiraptera, Thysanoptera. Melbourne: CSIRO Publishing, Australia.

Smithers, C.N. 1972. The classification and phylogeny of the Psocoptera. Memoirs of the Australian Museum 14: 1–349.

Yoshizawa, K., C. Lienhard, and K.P. Johnson. 2006. Molecular systematics of the suborder Trogiomorpha (Insecta: Psocodea: ‘Psocoptera’). Zoological Journal of the Linnean Society 146: 287–299.

About This Page

Emilie Bess
Illinois Natural History Survey, Champaign, Illinois, USA

Kevin P. Johnson
Illinois Natural History Survey, Champaign, Illinois, USA

Correspondence regarding this page should be directed to Emilie Bess at and Kevin P. Johnson at

All Rights Reserved.

Citing this page:

Bess, Emilie and Kevin P. Johnson. 2009. Psoquillidae. Version 18 March 2009 (under construction). http://tolweb.org/Psoquillidae/14452/2009.03.18 in The Tree of Life Web Project, http://tolweb.org/

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