Boloria
Clossiana , Proclossiana
Thomas Simonsen, Niklas Wahlberg, and Andrew V. Z. Brower
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close boxIntroduction
A holarctic genus. Many species are boreal or montane, with disjunct distributions on isolated mountain ranges.Characteristics
Putative autapomorphies for Boloria include (Simonsen 2005, 2006):- Uncus bifid
- Dorsal wall of bursa ostium/antrum raised vertically with a sclerotized, arched plate just posterior to the insertion of ductus seminalis
- Bursa ostium/antrum with sclerotized ventral plates
Putative synapomorphies for the subgenus Boloria and B. (Proclossiana) eunomia include (Simonsen 2005):
- Setae on uncus oriented towards the base of uncus
- Uncus with pronounced ventral ridges
- Distal margin of male valve clearly concave
Putative autapomorphies for the subgenus Boloria include (Simonsen 2005, 2006):
- Phallus with two posterior, dorso-lateral areas with strongly sclerotized microtrichia
- Each half of the male juxta in dorsal view outwards pointed spatulate with convex inner margin
Putative autapomorphies for subgenus Clossiana include (Simonsen 2005, 2006):
- A pronounced dent present dorsally where uncus and tegumen join
- A large, membranous centro-diagonal area present on the inner surface of the male valve
- Female abdominal venter 8 with two large, weakly sclerotized, scale covered latero-posterior areas
References
Simonsen, T. J. 2005. Boloria phylogeny (Lepidoptera: Nymphalidae): tentatively reconstructed on the basis of male and female genitalic morphology. Systematic Entomology 30:653-665.
Simonsen, T. J. 2006. Fritillary phylogeny, classification and larval hostplants: reconstructed mainly on the basis of male and female genitalic morphology (Lepidoptera: Nymphalidae: Argynnini). Biological Journal of the Linnean Society 89: 627-673.
Simonsen, T. J., N. Wahlberg, A. V. Z. Brower, and R. de Jong. 2006. Morphology, molecules and Fritillaries: approaching a stable phylogeny for Argynnini (Lepidoptera: Nymphalidae). Insect Systematics & Evolution 37: 405-418.
Title Illustrations
| Scientific Name | Boloria euphrosyne |
|---|---|
| Location | Gait Barrows NNR Lancashire, UK |
| Specimen Condition | Live Specimen |
| Source | Pearl-bordered Fritillary Bolaria euphrosyne |
| Source Collection | Flickr |
| ToL Image Use |
 This media file is licensed under the Creative Commons Attribution-NonCommercial-NoDerivs License - Version 2.0.
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| Copyright | © 2007 Graham |
| Scientific Name | Boloria napaea |
|---|---|
| Location | NW Russia |
| Specimen Condition | Dead Specimen |
| Identified By | Niklas Wahlberg |
| ToL Image Use |
This media file is licensed under the Creative Commons Attribution-NonCommercial License - Version 3.0.
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| Copyright |
© Niklas Wahlberg
|
About This Page
Thomas Simonsen
University of Alberta, Edmonton, Alberta, Canada
Niklas Wahlberg
University of Turku, Finland
Middle Tennessee State University, Murfreesboro, Tennessee, USA
Correspondence regarding this page should be directed to Thomas Simonsen at , Niklas Wahlberg at , and Andrew V. Z. Brower at
Page copyright © 2006 Thomas Simonsen, Niklas Wahlberg, and
All Rights Reserved.
- First online 04 October 2006
- Content changed 25 March 2007
Citing this page:
Simonsen, Thomas, Wahlberg, Niklas, and Brower, Andrew V. Z. 2007. Boloria . Clossiana , Proclossiana . Version 25 March 2007 (under construction). http://tolweb.org/Boloria/70413/2007.03.25 in The Tree of Life Web Project, http://tolweb.org/
